The aim of this course is to provide participants with an overview of morphometric approaches to studying morphological integration and modularity. The concepts of integration and modularity will be introduced and discussed in different contexts (e.g. development, individual variation, evolutionary change). The theoretical basis and application of different methods for analyzing integration and modularity in geometric morphometric data will be presented. Lectures will be combined with hands-on demonstrations of the analyses. Participants are encouraged to bring their own morphometric data for analysis and discussion in the course.
Prof. Chris Klingenberg
(University of Manchester, United Kingdom).
Dr. Soledad De Esteban-Trivigno
(Transmitting Science, Spain).
Graduate or postgraduate degree in Biomedical, Life or Earth Sciences, basic knowledge of statistics and personal computers. Knowledge of Geometric Morphometrics. All participants must bring their own personal laptop. We would like to encourage participants to bring along their own morphometrics data, in case you have it.
1. Brief review of geometric morphometrics: Procrustes fit, shape space, covariance matrices, etc.
2. PCA for characterizing patterns of integration.
3. Partial least squares and integration.
4. Practice: patterns of integration.
5. Indices of integration.
6. Modularity: concepts and analysis.
7. Modularity hypotheses: where do they come from?
8. Practice: Indices, allometry and comparisons.
9. Comparing patterns of integration.
10. Developmental integration and modularity.
11. Evolutionary integration and modularity.
12. Practice: integration & modularity.
13. Allometry as a factor of integration.
14. Multilevel analyses of integration and modularity.
15. Presentations of group work.
- Klingenberg CP, Zaklan SD (2000) Morphological integration between developmental compartments in the Drosophila wing. Evolution, 54: 1273-1285.
- Klingenberg CP, Badyaev AV, Sowry SM, Beckwith NJ (2001) Inferring developmental modularity from morphological integration: Analysis of individual variation and asymmetry in bumblebee wings. American Naturalist, 157: 11-23.
- Klingenberg CP (2003) Developmental instability as a research tool: Using patterns of fluctuating asymmetry to infer the developmental origins of morphological integration, Pages 427-442 in Polak M, ed. Developmental instability: Causes and consequences. New York, Oxford University Press.
- Klingenberg CP, Mebus K, Auffray J-C (2003) Developmental integration in a complex morphological structure: How distinct are the modules in the mouse mandible?. Evolution & Development, 5: 522-531.
- Klingenberg CP (2004) Integration, modules and development: Molecules to morphology to evolution, Pages 213-230 in Pigliucci M, Preston K, eds. Phenotypic integration: Studying the ecology and evolution of complex phenotypes. New York, Oxford University Press.
- Klingenberg CP (2005) Developmental constraints, modules and evolvability, Pages 219-247 in Hallgrímsson B, Hall BK, eds. Variation. San Diego, Academic Press.
- Breuker CJ, Gibbs M, Van Dyck H, Brakefield PM, Klingenberg CP, Van Dongen S (2007) Integration of wings and their eyespots in the speckled wood butterfly Pararge aegeria. Journal of Experimental Zoology (Molecular and Developmental Evolution), 308B: 454-463.
- Klingenberg CP (2008) Morphological integration and developmental modularity. Annual Review of Ecology, Evolution and Systematics, 39: 115-132.
- Klingenberg CP (2009) Morphometric integration and modularity in configurations of landmarks: Tools for evaluating a-priori hypotheses. Evolution & Development, 11: 405-421.
- Singh N, Harvati K, Hublin J-J, Klingenberg CP (2012) Morphological evolution through integration: A quantitative study of cranial integration in Homo, Pan, Gorilla and Pongo. Journal of Human Evolution, 62: 155-164.
- Klingenberg CP, Duttke S, Whelan S, Kim M (2012) Developmental plasticity, morphological variation and evolvability: A multilevel analysis of morphometric integration in the shape of compound leaves. Journal of Evolutionary Biology, 25: 115-129.
28 hours on-site.
|Monday 27th January 2014:
18:00: Meeting point in Plaza Catalunya (Barcelona) to take the bus to Els Hostalets de Pierola. If you are planning to arrive later, you can find more information in How to arrive.
8:30 to 9:00: Breakfast
8:30 to 9:00: Breakfast
|Thursday 30th January 2014:
8:30 to 9:00: Breakfast
8:30 to 9:00: Breakfast.
You will find below some testimonials from former participants to previous editions of this course:
- “In my point of view the course was quite comprehensive and well structured.”
Participants’ comment in the course evaluation (1st Edition).
Places are limited to 20 participants and will be occupied by strict registration order.
This course can be validated by 1.5 ECTS (European Credit Transfer System) by the Life Science Zurich Graduate School, Zurich, Switzerland.
Only Lunches Included option (no accommodation, no dinners):
Reduced registration fee until September 30, 2013: 375 €. Full registration fee after September 30, 2013: 525 €. Reduced registration fee extended to November 30, 2013.
This option only includes course material, coffee breaks and lunches. Please be aware that from Barcelona centre it takes one hour by car and one hour and a half by public transport to arrive to Els Hostalets de Pierola.
All Inclusive option:
Reduced registration fee until September 30, 2013: 585 € (375 € course fee + 210 € All Inclusive option). Full registration fee after September 30, 2013: 735 € (525 € course fee + 210 € All Inclusive option). Reduced registration fee extended to November 30, 2013.
Participation fees are all inclusive, and include transport from Barcelona city, course material, lodging, breakfasts, coffee breaks, lunches, dinners and welcome party.
Participants in any editions of this course are entitled to a 20 % discount on the course fee of the incoming course Quantitative Genetics of Shape - 2nd Edition.
Former participants will have a 5 % discount on the current course fee.
We offer the possibility of paying in two instalments (contact us at email@example.com).
Please complete and submit your Registration Form (see below); we will confirm your acceptance by e-mail.
If you wish to cancel your participation in this course, cancellations up to 20 days before the course start date will incur a 30 % cancellation fee. For later cancellations, or non-attendance, the full course fee will be charged.
If Transmitting Science must cancel this course due to unforeseen circumstances beyond the control of Transmitting Science, you will either be entitled to a full refund of the course fee, or your fee can be credited toward a future course / workshop. Transmitting Science is not responsible for travel fees, or any expenses incurred by you as a result of such cancellation. Every effort will be made to avoid the cancellation of any planned course / workshop.
The course will take place in the town of Els Hostalets de Pierola, Barcelona (Spain).
- Only Lunches Included option (no accommodation, no dinners):
In that case there are different options for accommodation, all of them in places other than Els Hostalets de Pierola. You may stay in: Barcelona city, Martorell or Igualada. Information about Hotels in Martorell here. Information about Hotels in Igualada here.
How to arrive to Els Hostalets de Pierola from Barcelona city, Martorell and Igualada
- All Inclusive option:
To make it affordable, you will be hosted in the Youth Hostel Can Roviralta, in four-bed dormitories (common bathrooms), although you will only share rooms with other classmates. The possibility exists that the dormitories will be mixed. Therefore, if this is an inconvenience for you, and you require staying in a dormitory with only women, or only men, please state it in the “Comments” field of the Registration Form (see below).
There are no Hotels in Els Hostalets de Pierola. However, if you do not want to share rooms, but you want to apply for the All Inclusive option staying in Els Hostalets de Pierola, please contact us at firstname.lastname@example.org.
How to arrive to Els Hostalets de Pierola.
Unfortunately there are no internal grants available for this course. However some discounts are offered to people belonging to some of the Associations, Institutions or Universities listed here. If you want to apply to this discount please state the name of your Association, Institution or University in the “Comments” field of the Registration Form (proof will be asked later). For information on External Financial Support, please check the link.
Spanish unemployed scientists, as well as Spanish PhD students without any grant to develop their PhD, could benefit from a 40 % discount on the course fee. If you want to ask for this discount, please contact us at email@example.com. That would apply for a maximum of 4 places and they will be covered by strict inscription order.
For further information contact: firstname.lastname@example.org.